:root {
  color-scheme: light;
  --ink: #16211f;
  --muted: #61706c;
  --line: #dce5e1;
  --paper: #f7faf8;
  --white: #ffffff;
  --green: #0c594d;
  --green-2: #1a7669;
  --blue: #255b8c;
  --gold: #caa45d;
  --shadow: 0 20px 55px rgba(18, 39, 37, 0.12);
  --radius: 8px;
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  font-family: "Inter", Arial, sans-serif;
  color: var(--ink);
  background: var(--paper);
  line-height: 1.6;
}

a {
  color: inherit;
}

.site-header {
  position: sticky;
  top: 0;
  z-index: 20;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  min-height: 76px;
  padding: 14px clamp(20px, 4vw, 64px);
  background: rgba(247, 250, 248, 0.92);
  border-bottom: 1px solid rgba(220, 229, 225, 0.9);
  backdrop-filter: blur(18px);
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 12px;
  text-decoration: none;
}

.brand-mark {
  display: grid;
  width: 42px;
  height: 42px;
  place-items: center;
  border-radius: 50%;
  color: var(--white);
  background: linear-gradient(135deg, var(--green), var(--blue));
  font-weight: 800;
}

.brand strong,
.brand small {
  display: block;
}

.brand strong {
  font-size: 1rem;
  line-height: 1.1;
}

.brand small {
  color: var(--muted);
  font-size: 0.76rem;
}

.site-nav {
  display: flex;
  align-items: center;
  gap: clamp(14px, 2vw, 28px);
  color: #31413d;
  font-size: 0.92rem;
  font-weight: 650;
}

.site-nav a {
  text-decoration: none;
}

.site-nav a:hover {
  color: var(--green-2);
}

.nav-toggle {
  display: none;
  width: 42px;
  height: 42px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--white);
}

.nav-toggle span {
  display: block;
  width: 18px;
  height: 2px;
  margin: 4px auto;
  background: var(--ink);
}

.hero {
  position: relative;
  min-height: calc(100vh - 76px);
  display: flex;
  align-items: center;
  overflow: hidden;
  color: var(--white);
  background: #0d1516;
}

.hero-image,
.hero-overlay {
  position: absolute;
  inset: 0;
}

.hero-image {
  width: 100%;
  height: 100%;
  object-fit: cover;
}

.hero-overlay {
  background:
    linear-gradient(90deg, rgba(7, 18, 18, 0.92) 0%, rgba(7, 18, 18, 0.76) 38%, rgba(7, 18, 18, 0.26) 100%),
    linear-gradient(0deg, rgba(7, 18, 18, 0.45), rgba(7, 18, 18, 0));
}

.hero-content {
  position: relative;
  z-index: 1;
  width: min(740px, calc(100% - 40px));
  margin-left: clamp(20px, 7vw, 96px);
  padding: 80px 0 110px;
}

.eyebrow {
  margin: 0 0 12px;
  color: var(--gold);
  font-size: 0.75rem;
  font-weight: 800;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}

h1,
h2,
h3,
p {
  overflow-wrap: break-word;
}

h1,
h2 {
  margin: 0;
  font-family: "Source Serif 4", Georgia, serif;
  line-height: 1.04;
}

h1 {
  max-width: 720px;
  font-size: clamp(3.1rem, 6vw, 6.8rem);
}

h2 {
  font-size: clamp(2.2rem, 4vw, 4.5rem);
}

h3 {
  margin: 0;
  font-size: 1.08rem;
  line-height: 1.25;
}

.hero-copy {
  max-width: 620px;
  margin: 24px 0 0;
  color: rgba(255, 255, 255, 0.86);
  font-size: clamp(1.04rem, 1.4vw, 1.28rem);
}

.hero-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 12px;
  margin-top: 34px;
}

.button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 48px;
  padding: 12px 18px;
  border: 1px solid transparent;
  border-radius: var(--radius);
  font-weight: 800;
  text-decoration: none;
}

.button.primary {
  color: var(--white);
  background: var(--green-2);
}

.button.primary:hover {
  background: var(--green);
}

.button.secondary {
  color: var(--white);
  border-color: rgba(255, 255, 255, 0.44);
  background: rgba(255, 255, 255, 0.08);
}

.button.secondary:hover {
  background: rgba(255, 255, 255, 0.16);
}

.band {
  background: var(--white);
  border-bottom: 1px solid var(--line);
}

.intro {
  padding: clamp(46px, 7vw, 88px) clamp(20px, 5vw, 72px);
}

.top-contact {
  padding: 30px clamp(20px, 5vw, 72px);
}

.top-contact-inner {
  display: grid;
  grid-template-columns: minmax(180px, 0.65fr) minmax(280px, 1.35fr);
  align-items: center;
  gap: clamp(20px, 4vw, 60px);
  max-width: 1180px;
  margin: 0 auto;
}

.top-contact h2 {
  font-size: clamp(1.75rem, 3vw, 3rem);
}

.contact-links {
  display: flex;
  flex-wrap: wrap;
  gap: 10px;
}

.contact-links,
.social-links {
  grid-column: 2;
}

.contact-links a {
  display: inline-flex;
  align-items: center;
  min-height: 42px;
  padding: 9px 13px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  color: #31413d;
  background: var(--paper);
  font-weight: 750;
  text-decoration: none;
}

.contact-links a:hover,
.contact-links a:focus-visible {
  border-color: rgba(20, 117, 107, 0.42);
  color: var(--green);
  background: #ffffff;
}

.social-links {
  display: flex;
  flex-wrap: wrap;
  gap: 10px;
  margin-top: -8px;
}

.social-links a {
  display: inline-grid;
  width: 42px;
  height: 42px;
  place-items: center;
  border: 1px solid var(--line);
  border-radius: 50%;
  color: var(--green);
  background: var(--paper);
  text-decoration: none;
}

.social-links a:hover,
.social-links a:focus-visible {
  border-color: rgba(20, 117, 107, 0.42);
  color: var(--white);
  background: var(--green);
}

.social-links svg {
  width: 21px;
  height: 21px;
  fill: currentColor;
}

.section-grid {
  display: grid;
  grid-template-columns: minmax(220px, 0.9fr) minmax(280px, 1.25fr);
  gap: clamp(28px, 5vw, 80px);
  max-width: 1180px;
  margin: 0 auto;
}

.section-grid p:last-child {
  margin: 0;
  color: #31413d;
  font-size: clamp(1.05rem, 1.6vw, 1.35rem);
}

.rna-wordmark {
  padding: clamp(34px, 5vw, 70px) clamp(20px, 5vw, 72px);
  background: #fbfcfb;
  border-bottom: 1px solid var(--line);
}

.rna-wordmark img {
  display: block;
  width: min(100%, 1040px);
  margin: 0 auto;
  border-radius: var(--radius);
}

.section {
  max-width: 1180px;
  margin: 0 auto;
  padding: clamp(60px, 8vw, 110px) clamp(20px, 5vw, 40px);
}

.section-heading {
  max-width: 760px;
  margin-bottom: 36px;
}

.research-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 16px;
}

.research-card,
.publication-card,
.person {
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--white);
  box-shadow: 0 12px 35px rgba(18, 39, 37, 0.06);
}

.research-card {
  min-height: 290px;
  padding: 24px;
}

.card-index,
.pub-year,
time {
  color: var(--blue);
  font-size: 0.78rem;
  font-weight: 800;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.research-card h3 {
  margin-top: 48px;
}

.research-card p,
.publication-list p,
.person p,
.timeline p,
.join-section p,
.site-footer p {
  color: var(--muted);
}

.split-section {
  display: grid;
  grid-template-columns: minmax(240px, 0.6fr) minmax(280px, 1fr);
  gap: clamp(28px, 6vw, 80px);
}

.team-list {
  display: grid;
  gap: 14px;
}

.team-groups {
  display: grid;
  gap: 34px;
}

.team-subheading {
  margin: 0 0 14px;
  color: var(--green);
  font-size: 0.86rem;
  font-weight: 800;
  letter-spacing: 0.1em;
  text-transform: uppercase;
}

.person {
  display: flex;
  align-items: center;
  gap: 16px;
  padding: 18px;
}

.person-photo {
  flex: 0 0 auto;
  width: 76px;
  height: 76px;
  border: 3px solid #edf4f1;
  border-radius: 50%;
  object-fit: cover;
}

.avatar {
  flex: 0 0 auto;
  display: grid;
  width: 58px;
  height: 58px;
  place-items: center;
  border-radius: 50%;
  color: var(--white);
  background: var(--green);
  font-weight: 800;
}

.open-role {
  border-color: rgba(202, 164, 93, 0.7);
}

.open-role .avatar {
  background: var(--gold);
}

.former-members .person {
  background: #fbfcfb;
}

.former-members .person-photo {
  filter: saturate(0.86);
}

.person p {
  margin: 4px 0 0;
}

.publication-list {
  display: grid;
  grid-auto-columns: calc((100% - 48px) / 4);
  grid-auto-flow: column;
  gap: 16px;
  overflow-x: auto;
  padding-bottom: 14px;
  scroll-snap-type: x mandatory;
  scrollbar-color: var(--blue) rgba(17, 71, 67, 0.12);
}

.publication-card {
  color: inherit;
  display: block;
  min-height: 100%;
  padding: 24px;
  scroll-snap-align: start;
  text-decoration: none;
  transition: border-color 180ms ease, box-shadow 180ms ease, transform 180ms ease;
}

.publication-card:hover,
.publication-card:focus-visible {
  border-color: rgba(20, 117, 107, 0.48);
  box-shadow: 0 18px 42px rgba(18, 39, 37, 0.12);
  transform: translateY(-2px);
}

.publication-card:focus-visible {
  outline: 3px solid rgba(20, 117, 107, 0.24);
  outline-offset: 3px;
}

.publication-list h3 {
  margin: 10px 0 8px;
}

.pub-year {
  margin: 0;
}

.timeline {
  border-left: 2px solid var(--line);
  display: grid;
  gap: 28px;
  margin-left: 10px;
  padding-left: 28px;
}

.timeline article {
  position: relative;
}

.timeline article::before {
  content: "";
  position: absolute;
  top: 6px;
  left: -36px;
  width: 12px;
  height: 12px;
  border: 3px solid var(--paper);
  border-radius: 50%;
  background: var(--green-2);
  box-shadow: 0 0 0 1px var(--green-2);
}

.timeline p {
  margin: 8px 0 0;
  max-width: 820px;
}

.join-section {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 28px;
  margin: 0 clamp(20px, 5vw, 72px) clamp(56px, 8vw, 100px);
  padding: clamp(30px, 5vw, 56px);
  border-radius: var(--radius);
  color: var(--white);
  background: linear-gradient(135deg, var(--green), #173d5f);
  box-shadow: var(--shadow);
}

.join-section h2 {
  max-width: 760px;
}

.join-section p {
  max-width: 770px;
  color: rgba(255, 255, 255, 0.82);
}

.site-footer {
  display: grid;
  grid-template-columns: minmax(240px, 0.8fr) minmax(320px, 1.2fr);
  align-items: stretch;
  gap: 28px;
  padding: 40px clamp(20px, 5vw, 72px);
  color: var(--white);
  background: #101817;
}

.site-footer h2 {
  font-size: 1.7rem;
}

.site-footer address {
  color: rgba(255, 255, 255, 0.78);
  font-style: normal;
}

.site-footer a {
  color: var(--white);
}

.footer-contact {
  display: grid;
  align-content: center;
  gap: 16px;
}

.map-frame {
  min-height: 260px;
  overflow: hidden;
  border: 1px solid rgba(255, 255, 255, 0.14);
  border-radius: var(--radius);
  background: #1b2524;
}

.map-frame iframe {
  display: block;
  width: 100%;
  height: 100%;
  min-height: 260px;
  border: 0;
  filter: grayscale(0.15) contrast(0.95);
}

.lab-cat {
  position: fixed;
  right: clamp(14px, 3vw, 36px);
  bottom: 14px;
  z-index: 18;
  width: clamp(94px, 12vw, 138px);
  pointer-events: none;
}

.lab-cat svg {
  display: block;
  width: 100%;
  filter: drop-shadow(0 10px 18px rgba(18, 39, 37, 0.18));
}

.cat-tail {
  animation: cat-tail-wag 1.8s ease-in-out infinite;
  transform-box: fill-box;
  transform-origin: 14% 80%;
}

.cat-eye {
  transform-box: fill-box;
  transform-origin: center;
}

.cat-lid {
  opacity: 0;
  transform: scaleY(0);
  transform-box: fill-box;
  transform-origin: center top;
}

.lab-cat.is-blinking .cat-lid {
  animation: cat-click-blink 260ms ease-in-out;
}

.cat-pupil {
  transform: translate(var(--eye-x), var(--eye-y));
  transition: transform 120ms ease-out;
}

@keyframes cat-tail-wag {
  0%,
  100% {
    transform: rotate(-4deg);
  }

  50% {
    transform: rotate(8deg);
  }
}

@keyframes cat-click-blink {
  0%,
  100% {
    opacity: 0;
    transform: scaleY(0);
  }

  50% {
    opacity: 1;
    transform: scaleY(1);
  }
}

@media (max-width: 980px) {
  .research-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .publication-list {
    grid-auto-columns: calc((100% - 16px) / 2);
  }

  .top-contact-inner,
  .split-section,
  .section-grid {
    grid-template-columns: 1fr;
  }

  .contact-links,
  .social-links {
    grid-column: 1;
  }
}

@media (max-width: 760px) {
  .site-header {
    min-height: 68px;
  }

  .nav-toggle {
    display: block;
  }

  .site-nav {
    position: absolute;
    top: 68px;
    right: 16px;
    left: 16px;
    display: none;
    flex-direction: column;
    align-items: stretch;
    gap: 0;
    padding: 10px;
    border: 1px solid var(--line);
    border-radius: var(--radius);
    background: var(--white);
    box-shadow: var(--shadow);
  }

  .site-nav.open {
    display: flex;
  }

  .site-nav a {
    padding: 12px;
  }

  .hero {
    min-height: 720px;
  }

  .hero-overlay {
    background: linear-gradient(90deg, rgba(7, 18, 18, 0.95), rgba(7, 18, 18, 0.55));
  }

  .hero-content {
    width: calc(100% - 40px);
    margin: 0 auto;
  }

  .research-grid {
    grid-template-columns: 1fr;
  }

  .publication-list {
    grid-auto-columns: minmax(260px, 86%);
  }

  .join-section {
    flex-direction: column;
    align-items: flex-start;
  }

  .site-footer {
    grid-template-columns: 1fr;
  }

  .lab-cat {
    width: 86px;
    opacity: 0.92;
  }
}

@media (max-width: 430px) {
  h1 {
    font-size: 2.75rem;
  }

  .button {
    width: 100%;
  }

  .person {
    align-items: flex-start;
  }

  .lab-cat {
    display: none;
  }
}

@media (prefers-reduced-motion: reduce) {
  .lab-cat.is-blinking .cat-lid,
  .cat-tail {
    animation: none;
  }
}